Significator
A significator is the planet a chart assigns to stand in for one of the parties in the question — you, the thing asked about (the tradition calls that the quesited), or a third person. It's a role a planet holds, not a quality it has.
The fuller picture
Assigning a significator is a lookup, not a judgement call. Decide which house governs the matter, read the sign on that house's cusp, and the planet ruling that sign is the significator of the matter. The querent — the person asking — takes the ruler of the first house, Lord 1, by the same route, and the Moon assists throughout. Because it's a role rather than a property, one planet can be read twice in a single chart and mean two different things. In one worked chart the same Mars, at the same degree with the same dignity, describes a sober and realistic sense of what's possible when it's read as the querent, and ample internal resource for the goal when it's read as the thing wanted. Nothing about the planet changed — only the role did.
Only seven bodies rule anything in this tradition: Saturn, Jupiter, Mars, the Sun, Venus, Mercury and the Moon. Uranus, Neptune and Pluto never rule a sign or a house, so they're never significators — which isn't the same as being ignored. They appear as witnesses, describing a matter they have no authority over, most often when one of them rises or throws a hard aspect at the querent's ruler or the Moon.
Seven rulers across twelve houses means one planet routinely holds more than one role, and that collision is information rather than a problem. When two topics land on the same planet, they're read as entangled — a planet standing for a brother that also rules the tenth house carries the brother and his business at once. When the planet standing for the person asking is also the planet standing for the thing asked about, something structurally different happens: there's no contact left to wait for, because the two parties are one body, and the identity itself is what says they're connected. That case is treated on the querent page.
Two significators are rarely the whole cast. Some things carry a general significator regardless of which house governs them — Venus for children, the Sun for the man in a question where the topic has already claimed Venus. Anyone the question reaches through somebody else takes a derived significator: a house standing for a person becomes that person's own first house, and the count continues from there. Several familiar house meanings are already the result of exactly that count — the eighth is a partner's money because it's the second house from the seventh, the ninth is in-laws because it's the third from the seventh. There's also a dedicated step for the rulerships that bear on the matter without being the principal pair, and an intercepted sign hands its house a second ruler, which is how the co-ruler of an illness can turn out to be sitting in somebody else's house of health.
The order in which all this is examined is itself doctrine, and it's deliberately slower than reading a chart intuitively: identify the querent's planet and weigh its essential and accidental condition, then the planet standing for the thing asked about, weighed the same way, then the Moon — three actors, each weighed twice — and only then look at the aspects between them, and only after that reach a verdict. The rigidity is the point; it stops a student seizing on whichever feature of the chart happens to catch their eye. One caution belongs alongside it. Which house a significator counts as occupying can turn on a few arcminutes, because a planet within five degrees of the next cusp, and in the same sign as that cusp, is read as belonging to the house it's about to enter rather than the one it's technically still in.
In your chart
iHorary names every significator on the wheel by its role and its lordship together — "Querent · L1" sitting beside "Money · L2" as an equal — and when one planet holds two roles it gets a single plate carrying both labels rather than appearing twice. The Moon joins as co-significator only if no other role has already claimed her. The judgment layer then runs its per-planet condition checks over exactly the two principals, and the Moon is assessed on her own terms.
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